In examples of chemically or genetically induced tumor formation, an increased rate of aneuploidaneuploidy acts as a more effective inhibitor than initiator of tumorigenesis. This reveals a tumor-suppressive role for chromosomal instability in contexts where tumorigenesis is driven by chemical carcinogens or genetic alterations. [@weaver_aneuploidy_2007]

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Multiple studies establish that elevated aneuploidy paradoxically suppresses tumor formation in chemically and genetically induced cancer models, even though it promotes spontaneous tumors in aged animals, revealing context-dependent oncogenic and tumor-suppressive roles for chromosomal instability. The mechanistic basis for this dual behavior appears to involve aneuploidy-induced cellular stress that initially inhibits transformation, though aneuploid cells that do become malignant evolve specific adaptations—particularly Stat1 inactivation combined with Myc activation—to suppress immune infiltration and overcome the proliferative constraints normally imposed by chromosome missegregation. What remains contested is whether the tumor-suppressive effect of high aneuploidy in induced models reflects intrinsic proliferation defects that cannot be overcome, or whether sufficient selective pressure and time would eventually yield transformed clones with the necessary compensatory mutations, as the requirement for additional phenotypic changes beyond missegregation alone suggests that viable aneuploid tumor cells represent rare evolutionary solutions rather than inevitable outcomes.

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