Chromosomally unstable tumors exhibit increased immune cell infiltration compared to chromosomally stable tumors. This enhanced immune surveillance represents a fitness cost of chromosomal instability that limits tumor growth and malignancy in vivo. [@schubert_cancer_2021]

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Chromosomal instability is established to promote immune cell infiltration into tumors, creating an immunogenic environment that exposes chromosomally unstable cancer cells to immune surveillance. This infiltration occurs because the genomic chaos generated by ongoing chromosome missegregation produces heterogeneous neoantigens and stress signals that attract immune cells, as demonstrated both in mouse models and human cancers. However, for chromosomally unstable tumors to achieve malignancy, they must actively suppress this immune response through a conserved mechanism involving Stat1 signaling inactivation combined with Myc activation, which specifically alleviates the chromosomal instability-induced immune infiltration and allows aneuploid cells to persist despite their fitness costs. While the initial immunogenic consequences of chromosomal instability and the subsequent requirement for immune evasion are well-established, the precise molecular signals that trigger initial immune cell recruitment and whether all forms of chromosomal instability generate equally immunogenic environments remain areas requiring further investigation.

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